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In evolutionary biology, the term is also applied to males who are unwittingly investing parental effort in offspring that are not genetically their own.

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One often-overlooked subtlety of the word is that it implies that the husband is deceived, that he is unaware of his wife's unfaithfulness and may not know until the arrival or growth of a child plainly not his (as with cuckoo birds). A related word, first appearing in 1520, is wittol, which substitutes wit (in the sense of knowing) for the first part of the word, referring to a man aware of and reconciled to his wife's infidelity.

In Western traditions, cuckolds have sometimes been described as "wearing the horns of a cuckold" or just "wearing the horns".

The association is common in medieval folklore, literature, and iconography.

English usage first appears about 1250 in the satirical and polemical poem "The Owl and the Nightingale" (l. The term was clearly regarded as embarrassingly direct, as evident in John Lydgate's "Fall of Princes" (c. In the late 14th century, the term also appeared in Geoffrey Chaucer's "The Miller's Tale".

As noted above, the term cuckold is derivative of the mis-directed parental investment of birds who direct parental investment to the eggs that cuckoo birds have laid in their nests.

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